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40d:Tower-cap
Tower caps are a type of tree which only grow underground (subterranean). Once fully grown (which takes 3 years?) they can be designated for wood cutting like other trees, and produce Tower Cap logs.
Tree Farms
In is possible for tower caps to spontaneously grow in muddied subterranean areas only if you have discovered an underground river or underground pond. Unfortunately this is currently extremely difficult to do.
On the subject of spawning Tower Caps, Toady said:
Right now, you can't plant towercaps or bring spawn along, so you need to find a cave river or cave pool, in which case they will start growing underground if it's all working. It's trickier to find such things, so it's mostly impossible to do. Later, you should be able to plant them. There are no arbitrary vegetation caps now.
The "arbitary vegetation caps" comment is referring to the old limit of 200 trees growing by the cave river in the mud.